Natural selection causes adaptation, the fit between an organism and its environment. For example, the white and grey coloration of snowy owls living and breeding around the Arctic Circle provides camouflage from both predators and prey. In this Element, we explore a variety of such outcomes of the evolutionary process, including both adaptations and alternatives to adaptations, such as nonadaptive traits inherited from ancestors. We also explore how the concept of adaptation is used in evolutionary psychology and in animal behavior, (...) and the adequacy of methods used to confirm evolutionary accounts of human traits and behaviors. (shrink)

We address the controversy in the literature concerning the definition of holobionts and the apparent constraints on their evolution using concepts from community population genetics. The genetics of holobionts, consisting of a host and diverse microbial symbionts, has been neglected in many discussions of the topic, and, where it has been discussed, a gene-centric, species-centric view, based in genomic conflict, has been predominant. Because coevolution takes place between traits or genes in two or more species and not, strictly speaking, between (...) species, it may affect some traits but not others in either host or symbiont. Moreover, when interacting species pairs are embedded in a larger community, indirect ecological effects can alter the expected pairwise dynamics. Mode of symbiont transmission and the degree of host inbreeding both affect the extent of microbial mixing across host lineages and thereby the degree to which selection on one trait of either partner affects other aspects of a holobiont phenotype. We discuss several potential defining criteria for holobionts using community genetics and population genetics models, suggesting their application and limitations. Using community genetics models, we show how conflict between genomes can be self-limiting, while cooperation and mutualism tend to be self-accelerating. It is likely that this bias in the evolutionary dynamics of interaction between hosts and symbionts is an important feature of holobionts. This bias in the evolutionary dynamic could contribute to explaining the absence of cheaters from natural mutualisms, although cheaters are predicted by gene-centered conflict theory to cause the evolutionary instability of mutualisms. Additionally, it may help explain the more frequent origin of mutualisms from parasitic than from free-living systems, an evolutionary trajectory opposite to that predicted by genome conflict theory. (shrink)

Modern science’s ability to produce, store, and analyze big datasets is changing the way that scientific research is practiced. Philosophers have only begun to comprehend the changed nature of scientific reasoning in this age of “big data.” We analyze data-focused practices in biology and climate modeling, identifying distinct species of data-centric science: phenomena-laden in biology and phenomena-agnostic in climate modeling, each better suited for its own domain of application, though each entail trade-offs. We argue that data-centric practices in science are (...) not monolithic because the opportunities and challenges presented by big data vary across scientific domains. (shrink)

In this paper we consider some questions surrounding whether or not regional climate models “add value,” a controversial issue in climate science today. We highlight some objections frequently made about regional climate models both within and outside the community of modelers, including several claims that regional climate models do not “add value.” We show that there are a number of issues involved in the latter claims, the primary ones centering on the fact that different research questions are being pursued by (...) the modelers making the complaints against regional climate models. Further issues focus on historical deficiencies of particular—but not generalizable—failures of individual regional models. We provide tools to sort out these different research questions and particular failures, and to improve communication and understanding surrounding added value in climate modeling and philosophy of climate science. (shrink)

A specific form of research question, for instance, “What is the probability of a certain class of weather events, given global climate change, relative to a world without?” could be answered with the use of FAR or RR (Fraction of Attributable Risk or Risk Ratio) as the most common approaches to discover and ascribe extreme weather events. Kevin Trenberth et al. (2015) and Theodore Shepherd (2016) have expressed doubts in their latest works whether it is the most appropriate explanatory tool (...) or the way of public outreach concerning climate events and extremes. As an alternative, these researchers focus on complementary questions, for example, “How much did climate change affect the severity of a given storm?” advocating a “storyline” approach. New methods and new research questions are neither foreign, nor controversial from the standpoint of history and philosophy of science, especially those, related to public interest. Nevertheless, the new proposal has got a tepid reception from the majority of professionals of the given field. They argued that this approach can cause weakening of standards and neglecting of scientific method. The following paper attempts to find the roots of the supposed controversy. We claim inefficiency of uncompromising approach to D&A in absolute sense and assert that errors of a particular type may have a different level of concern in society, depending on the variety of contexts. Therefore, context defines the risk of over-estimation vs. under-estimation of harm. (shrink)

Despite the diversity of ageing, society and academics often describe and label older persons as a vulnerable group. As the term vulnerability is frequently interchangeably used with frailty, dependence, or loss of autonomy, a connection between older age and deficits is promoted. Concerning this, the question arises to what extent it may be helpful to refer to older persons as vulnerable specifically in the context of health care. After analyzing different notions of vulnerability, I argue that it is illegitimate to (...) conclude that older age is related to increased vulnerability. Much more, identifying older adults as a vulnerable group is closely related to ageism and can be associated with paternalistic benevolence and a tendency to overprotection, especially within health care. Additionally, even though older adults are more often in situations of increased vulnerability due to their potentially higher need for health care, I argue further that older adults mainly become a vulnerable group due to ageism. In this way, it can be concluded that the vulnerability of older adults does not originate in certain attributes of the group, but arises from a characteristic of society and, in turn, health personnel, namely ageism. Labeling older adults as vulnerable therefore is only helpful, when it is used to raise awareness of the widespread ageism in society, in this context, especially in the setting of health care, and the negative consequences thereof for older adults. (shrink)

As we advance into the twenty-first century, the evidence of climate change is all around us. In the introduction to this volume, we discuss some of the successes of climate science in understanding and attributing the causes of these changes, as well as some of the challenges it faces in addressing questions for which we do not yet have the answers. We focus on the role of climate models and the philosophical and conceptual problems facing climate modelers and climate modeling. (...) We then give the reader an outline of what they will find in each of the 15 Chapters in the volume. The book is divided into three major parts: Part 1 “Confirmation and Evidence”; Part 2 “Uncertainties and Robustness”; and Part 3 “Climate Models as Guides to Policy”. (shrink)

The conflation of two fundamentally distinct issues has generated serious confusion in the philosophical and biological literature concerning the units of selection. The questions of how a unit of selection is defined, theoretically, is rarely distinguished from the question of how to determine the empirical accuracy of claims--either specific or general--concerning which unit(s) are undergoing selection processes. In this paper, I begin by refining a definition of the unit of selection, first presented by William Wimsatt, that is grounded in the (...) structure of natural selection models. I then explore the implications of this definition for empirical evaluation of group selection models. I examine an objection to Wimsatt’s definition raised by Elliott Sober, and I conclude by indicating how the proposed refined definition can clarify which sorts of empirical evidence could be brought to bear on group selection controversies. (shrink)

In this brief chapter, Lloyd sets the stage for the following three papers, most centrally, Santer et al., which discusses whether the satellite data fit with climate models. Its target is a paper by Douglass et al., which claimed that satellite and weather balloon data showed that the climate models were wrong and could not be trusted. The Santer and Wigley “Fact Sheet” gives a nontechnical summary of what is wrong with the Douglass paper, while the full story is in (...) Chap. 5, a reprint of the Santer et al. paper. The context and import of this work is discussed in Lloyd. In this introduction, Lloyd emphasizes the timeliness of this work, as climate deniers testify in Congress presently. (shrink)

Every philosophical involvement with language centres on the notion of representation. There is controversy over what language represents. The answers can be classified and used as a basis for a systematic survey: 1. Language represents the world. 2. Language does not represent the world but our ideas of the world. 3. Language represents our ideas (of the world) badly. 4. Language not only represents badly; it does not represent anything. 5. Without language there would be no representation of a higher (...) order and thus no thought. The main intention behind this introduction to linguistic philosophy lies in revealing the underlying ordering principle for the many simultaneous and non-simultaneous attempts at a philosophy of language. (shrink)

In the context of a broad welcome to Bullot & Reber's (B&R's) proposals concerning the incorporation of contextual awareness into the study of the psychology of art appreciation, I raise two concerns. First, the proposal makes no allowance for degrees of relevance of contextual awareness to appreciation. Second, the authors assume that and can be maintained as separate phases or modes, but this may be more problematic than anticipated.