Abstract

I argue that Descartes explains physiology in terms of whole systems, and not in terms of the size, shape and motion of tiny corpuscles (corpuscular mechanics). It is a standard, entrenched view that Descartes’ proper means of explanation in the natural world is through strict reduction to corpuscular mechanics. This view is bolstered by a handful of corpuscular–mechanical explanations in Descartes’ physics, which have been taken to be representative of his treatment of all natural phenomena. However, Descartes’ explanations of the ‘principal parts’ of physiology do not follow the corpuscular–mechanical pattern. Des Chene has identified systems in Descartes’ account of physiology, but takes them ultimately to reduce down to the corpuscle level. I argue that they do not. Rather, Descartes maintains entire systems, with components selected from multiple levels of organization, in order to construct more complete explanations than corpuscular mechanics alone would allow.