This is an introduction to the Symposium on “Evolution, Original Sin, and the Fall,” which has been designed as a thematic section for Zygon: Journal of Religion and Science. The Symposium investigates the enduring question of whether hamartiology (the theological study of sin) is compatible with evolutionary theory. We trace the origins of this question to the debate between Modernists and Traditionalists at the turn of the previous century. Our contributors make headway in these discussions by delving into details, namely (...) by asking whether the hamartiological views of particular theologians, such as Augustine or Aquinas, can be reconciled with specific aspects of evolutionary theory. They also extend hamartiology in novel directions through the application of critical race theory and literary science to shed new light on the origin and transmission of original sin. (shrink)