Abstract

This work is an examination of the metaphysical presuppositions involved in the science of organic form. Taking the dichotomy of structuralism versus functionalism in nineteenth century biology as the central subject of my study, I explore a network of unquestioned premises and isolate areas where empirical research programs and underlying metaphysical commitments both inform and hinder each other. ;I begin with the Cuvier-Geoffroy debate of 1830--a debate that clearly articulates the tensions between structuralist and functionalist approaches to organic form. On the side of functionalism, I am led to confront the rich concept of teleological causality. The concept of teleology, frequently taken to be semantically univocal, in fact unfolds into a cluster of related notions. I begin to demarcate significant differences between "intentional," "heuristic," "cosmic," and "organismic" teleology. On the side of structuralism, I set out to unearth the presuppositions of materialist versus idealist conceptions of structure--showing where naturalists have been both liberated and enslaved by their respective metaphysical commitments. With the basic divisions between structuralist and functionalist drawn, I set out to trace the dichotomy through the paradigm shift from fixism to transformism. ;In Darwin's system the structural/functional tensions become synthesized into complementary principles of causality. Here I explore the underlying presuppositional shifts which enabled Darwin to harmonize the previously intractable oppositions. These shifts include: rethinking teleology without "design," rethinking the archetype as real progenitor, and adopting a pluralistic perspective regarding causal mechanisms. ;The relative success of Darwin's appropriation of structuralism and functionalism leads to a discussion of the dichotomous character of the central subject itself. I argue that at the root of the dichotomous characterization of structure and function lies an impoverished understanding of teleology. I then offer "organicist" teleology as a powerful alternative to the French-Anglo tradition of "intentional" teleology. Presupposing the alternative teleology forms a challenge to the way in which French-Anglo naturalists have understood structural and functional relations. Together with Darwin's causal pluralism, a richer notion of teleology provides a different set of metaphysical assumptions and leads to an alternative model for empirical research programs