What is optimized in an optimal path?

Behavioral and Brain Sciences 36 (5):566 - 566 (2013)
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Abstract

An animal confronts numerous challenges when constructing an optimal navigational route. Spatial representations used for path optimization are likely constrained by critical environmental factors that dictate which neural systems control navigation. Multiple coding schemes depend upon their ecological relevance for a particular species, particularly when dealing with the third, or vertical, dimension of space

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