Biological attention to evolution and animal life has primarily emphasized a filiative approach that, although important, overlooks crucial dimensions highlighted by an ecological approach to animal human societies. Increased attention to singular animals and critical scrutiny of the operating definitions of society and culture indicates that vast dimensions of this area have been overlooked and remain to be studied. It is particularly important to pursue the aspects of signification, meaning, individuation, and subjectivity. Attention to animal human societies, or to animal (...) cultures that develop in the heart of human cultures, shows that humans and animals often form extimate relations based on particular aspects of animal subjectivity. With certain species we share a dense form of intertwining built on natural, cultural, and biographical histories. Beyond that, an argument on biosemiotic grounds maintains that culture is intrinsic to the living. (shrink)